The story kicks off when a teenager's premonition of a deadly racecar crash spares his life and those of a few lucky others. But death continues to stalk those who escaped it.

The sub-zero heroes from the worldwide blockbusters “Ice Age” and “Ice Age: The Meltdown” are back, on an incredible adventure...for the ages. Scrat is still trying to nab the ever-elusive nut (while, maybe, finding true love); Manny and Ellie await the birth of their mini-mammoth, Sid the sloth creates his own makeshift family by hijacking some dinosaur eggs; and Diego the saber-toothed tiger wonders if he’s growing too “soft” hanging with his pals. On a mission to rescue the hapless Sid, the gang ventures into a mysterious underground world, where they have some close encounters with dinosaurs, battle flora fauna run amuck – and meet a relentless, one-eyed, dino-hunting weasel named Buck.